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The Whites (Lepidoptera: Pieridae) of Papua Indonesia

by Henk van Mastrigt

The Pieridae are generally known as "Whites" because of the dominantly white coloured species worldwide, but there also many yellow, green and even bright red and orange coloured species. Even the approximal total number of species in the world is a point of discussion among scientists but it is something between 1,000 and 1,200 species. Parsons (1999) sets the number of New Guinea species on 128 but in Papua alone we recorded 139 species so the total number is probably higher. The genus Delias is the most divers with many endemic and isolated species. It is probably one of the most popular butterfly groups among collectors and scientists. The beautiful coloured and patterned hindwing underside is amazingly divers and attractive. Some species, especially in the Coliadinae, migrate over great distances and therefore many species have a wide distribution.

This page is under constant construction. This means that at any time new species can be added. Please check this page now and then for more information.

The checklist is based on relevant literature and completed with collection data.

Listed Papua species: 143 [xx endemic species for Papua (not sorted out yet)]

Coliadinae

Catopsilia Hübner, 1819

pomona (Fabricius, 1775)
crocale Cramer, 1775
catilla Cramer, 1779

scylla (Linnaeus, 1763)
ssp. etesia (Hewitson, [1867])
moluccarum Fruhstorfer, 1910
scylloides Fruhstorfer, 1902

pyranthe (Linnaeus, 1758)

Eurema Hübner, 1819

hecabe (Linnaeus, 1758)
ssp. oeta (Fruhstorfer, 1910)

alitha (Felder, 1862)
ssp. novaguineensis Shirozu & Yata, 1982
papuana Shirozu & Yata, 1981 (nec Butler, 1898)

blanda (Boisduval, 1836)
ssp. sahara (Fruhstorfer, 1912)

puella (Boisduval, 1832)
ssp. diotima (Fruhstorfer, 1910)

Gandaca Moore, 1896

butyrosa (Butler, 1875)
ssp. aiguina Fruhstorfer, 1910

Pierinae

Leuciacria Rothschild & Jordan, 1905

acuta Rothschild & Jordan 1905

Elodina C. & R. Felder, 1865

andropis Butler, 1876
ssp. namatia Fruhstorfer, 1910

hypatia C. & R. Felder, 1865
mulliercula Fruhstorfer, 1910
pasarga Fruhstorfer, 1910
doearilla Van Eecke, 1915

umbratica Grose-Smith, 1889
sada Fruhstorfer, 1910

definita Joicey & Talbot, 1916

Saletara Distant, 1885

corinna (Wallace, 1867)
ssp. cycinna (Hewitson, 1868)
hastia Fruhstorfer, 1910

Appias Hübner, 1819

paulina (Cramer, 1777)
ssp. saina Grose-Smith, 1894
sosias Fruhstorfer, 1913

albina (Boisduval, 1836)
ssp. albina (Boisduval, 1836)

celestina (Boisduval, 1832)
ssp. galerus Fruhstorfer, 1910

ada (Stoll, 1781)
ssp. thasia (Fruhstorfer, 1901)
plotina Fruhstorfer, 1902

Pareronia Bingham, 1907

jobaea (Boisduval, 1832) [endemic]

chinki Joicey & Noakes, 1915 [endemic]

Cepora Billberg, 1820

abnormis (Wallace, 1867)
euryxantha Honrath, 1892

aspasia (Stoll, 1790) [endemic]

perimale (Donovan, 1805)
ssp. wallaceana (C. & R. Felder, 1865)

Delias Hübner, 1819 (For more Delias of the World, see Les Day)

belisama group (xvii)

aruna (Boisduval, 1832)
ssp. aruna (Boisduval, 1832)
ssp. rona Rothschild, 1898
ssp. inferna Butler, 1871

hyparete group (xx)

argenthona (Fabricius, 1793)
ssp. balli Hulstaert, 1923

euphemia Grose-Smith, 1894 [endemic]

biaka Joicey & Noakes, 1915 [endemic]

talboti Joicey & Noakes, 1915 [endemic]

bosnikiana Joicey & Noakes, 1915 [endemic]

mysis (Fabricius, 1775)
ssp. nemea Fruhstorfer, 1910 [endemic]

lara (Boisduval, 1836)
ssp. lara (Boisduval. 1836)
cruentata Butler, 1865
maforensis Rothschild, 1915
oisyme Fruhstorfer, 1910
ssp. hideyoae Nakano, 1995 [endemic]

dorimene group (xviii)

gabia (Boisduval, 1832)
ssp. gabia (Boisduval, 1832)
ssp. aurantimacula Joicey & Talbot, 1922
ssp. marinda Hulstaert, 1924
ssp. felsina Fruhstorfer, 1910

mavroneria Fruhstorfer, 1914
ssp. mavroneria Fruhstorfer, 1914
ssp. flavidior Rothschild, 1915

nysa group (ii)

pulla Talbot, 1937
ssp. pulla Talbot, 1937 [endemic]
ssp. nimivinye Van Mastrigt, 2001 [endemic]

dice (Vollenhoven, 1865)
ssp. dice (Vollenhoven, 1865) nigroapicalis Roepke, 1955
ssp. mitisana Strand, 1916
pictula Strand, 1916
leucana Strand, 1916
ssp. latoclavata Van Eecke, 1924
ssp. latimarginata Joicey & Talbot, 1922
ssp. fulvoflava Rothschild, 1915

maudei Joicey & Noakes, 1915 [endemic]

isse group (xix)

ennia Wallace, 1867
ssp. ennia Wallace, 1867
ssp. mysolensis Rothschild, 1915
ssp. iere Grose-Smith, 1900
ssp. multicolor Joicey & Noakes, 1915
ssp. jobiana (Oberthür, 1894)
ssp. oetakwensis Rothschild, 1915

parennia Roepke, 1955 [endemic]

enniana (Oberthür, 1880)
ssp. enniana (Oberthür, 1880)
ssp. obsoleta Rothschild, 1915
ssp. contracta Talbot, 1928
ssp. kapaura Rothschild, 1915
ssp. majoripuncta Joicey & Talbot, 1922
ssp. reducta Rothschild, 1915

geraldina group (v)
sagessa subgroup

sagessa Fruhstorfer, 1910
ssp. anjae Schröder, 1977
ssp. sagessa Fruhstorfer, 1910

abrophora Roepke, 1955
ssp. abrophora Roepke, 1955
ssp. bugebu Van Mastrigt, 1996
ssp. okbibab Van Mastrigt, 1996

sinak Van Mastrigt, 1990

geraldina Smith, 1894
ssp. vogelcopensis Yagishita, 1993
ssp. onin Yagishita, 2003
ssp. vaneechoudi Roepke, 1955
ssp. siderea Roepke, 1955
flavescens Roepke, 1955
ssp. masakoae Nakano, 1998

aroae-pheres subgroup

angiensis Talbot, 1928 [endemic]

yabensis Joicey & Talbot, 1922 [endemic]
brevifascia Joicey & Talbot, 1922

approximata Joicey & Talbot, 1922 [endemic]
rectimargo Joicey & Talbot, 1922

pheres Jordan, [1912] [endemic]

inopinata Lachlan, 2000
ssp. sibatanii Van Mastrigt, 2003

balimensis Roepke, 1955 [endemic]

subapicalis Orr & Sibatani, 1985
ssp. orri Van Mastrigt, 2003

kenricki Talbot, 1937 [endemic]
ochraceus Joicey & Noakes, 1915

daniensis Van Mastrigt, 2003 [endemic]

imitator Kenrick, 1911 [endemic]

jordani Kenrick, 1909 [endemic]

fascelis subgroup

fascelis Jordan, [1912]
ssp. citrona Joicey & Talbot, 1922
ssp. sagawaorum Morita, 1996
ssp. paniaia Schmitt, 1992
ssp. amungme Van Mastrigt, 1996
ssp. ibelana Roepke, 1955
ssp. korupun Van Mastrigt, 1996
ssp. fascelis Jordan, [1912]

dortheysi Van Mastrigt, 2002 [endemic]

cuningputi subgroup

oktanglap Van Mastrigt, 1990
ssp. nishiyamai Yagishita, 1993
ssp. oktanglap Van Mastrigt, 1990

langda Gerrits & Van Mastrigt, [1993] [endemic]
ssp. watlangku Gerrits & Van Mastrigt, [1993]
ssp. langda Gerrits & Van Mastrigt, [1993]

stresemani group (iv)

rileyi Joicey & Talbot, 1922 [endemic]
ssp. rileyi Joicey & Talbot, 1922
ssp. yofona Schröder & Treadaway, 1982 comb. nov.
ssp. nishizawai Van Mastrigt & Sibatani, 1991

hikarui Yagishita, 1993 (probably hybrid) [endemic]

lecerfi Joicey & Talbot, 1922 [endemic]

cyclosticha Roepke, 1955

microsticha Rothschild, 1904
ssp. flavopicta Jordan, [1912]
ssp. weja Van Mastrigt, 2006
ssp. microsticha Rothschild, 1904
albifascia Talbot, 1928

nigropunctata Jordan & Noakes, 1915 [endemic]

eichhorni group (vi)

carstensziana Rothschild, 1915
ssp. carstensziana Rothschild, 1915
alcicornis Roepke, 1955
ssp. starensis Lachlan, 2000

leucobalia Jordan, [1912]
ssp. distincta Rothschild, 1915
ssp. ericetorum ROEPKE, 1955

eichhorni? subgroup

heliophora Roepke, 1955

muliensis Morinaka, Van Mastrigt & Sinabati, 1991

germana Roepke, 1955

antara Roepke, 1955
ssp. antara Roepke, 1955
ssp. solana Morinaka & Nakazawa, 1997

frater Jordan, [1912]
ssp. far Schröder& Treadaway, 1982
ssp. soror Toxopeus, 1944
ssp. frater Jordan, [1912]

toxopei Roepke, 1955 [endemic]
ssp. morosa Roepke, 1955
ssp. uranoi Yagishita, 1993
ssp. toxopei Roepke, 1955
ssp. nipsan Van Mastrigt, 1996

catisa Jordan, [1912]
ssp. wisseliana Roepke, 1955
ssp. morinakai Yagishiya, 1993
ssp. sumbole Van Mastrigt, 2001
ssp. aurostriga Roepke, 1955
ssp. catisa Jordan, [1912]

kummeri group (x)

isocharis Rothschild & Joicey, 1907
ssp. latiapicalis Joicey & Talbot, 1922
paniaia Roepke, 1955

strix Yagishita, 1993
ssp. strix Yagishita, 1993 comb. nov.

ligata Rothschild, 1904
ssp. dealbata Talbot, 1928
ssp. weylandensis Joicey & Talbot, 1922
ssp. interpolata Roepke, 1955

kummeri Ribbe, 1900
ssp. similis Talbot, 1928
ssp. athena Yagishita, 2003
ssp. chiekoae Nakano, 1995
ssp. rouffaer Yagishita, 1993
ssp. fumosa Roepke, 1955

fojaensis Van Mastrigt, 2006 [endemic]

alepa Jordan, [1912] [endemic]
ssp. kunupiensis Joicey & Talbot, 1922
ssp. orthobasis Roepke, 1955
ssp. alepa Jordan, [1912]

dixeyi Kenrick, 1909 [endemic]
obsoleta Küppers, 1978

bothwelli Kenrick, 1909 [endemic]
latimarginata Küppers, 1978
intermedia Küppers, 1978
lutea Küppers, 1978
lutes Küppers, 1978

marguerita Joicey & Talbot, 1922 [endemic]

pseudomarguerita Gerrits & Van Mastrigt, [1993] [endemic]

phippsi Joicey & Talbot, 1922 [endemic]
ssp. phippsi Joicey & Talbot, 1922
ssp. wisseli Roepke, 1955
ssp. mulia Gerrits & Van Mastrigt, [1993]

iltis group (viii)

iltis Ribbe, 1900
ssp. sibil Van Mastrigt, 1996

luctuosa Jordan, [1912]
ssp. magoda Gerrits & Yagishita, 2000
ssp. gottsi Gerrits & Van Mastrigt, [1993]
ssp. archboldi Roepke, 1955
butyracea Roepke, 1955
ssp. kuning Van Mastrigt, 1990

raymondi Schröder & Treadaway, 1982 [endemic]
ssp. raymondi Schröder & Treadaway, 1982 stat.nov.
ssp. shirahatai Morita, 1996
ssp. ogawai Morita, 1996

callista Jordan, [1912]
ssp. callipulchra Gerrits & Van Mastrigt, [1993]
ssp. callipareia Roepke, 1955
luteola Roepke, 1955
ssp. callista Jordan, [1912]
ssp. porquiaensis Yagishita, 1993

bakeri Kenrick, 1909 [endemic]

mesoblema subgroup

arabuana Roepke, 1955 [endemic]
ssp. modioensis Van Mastrigt, [1988]
ssp. asawaorum Morita, 1996
ssp. arabuana Roepke, 1955

mesoblema Jordan, [1912] [endemic]

awongkor Van Mastrigt, [1988]
ssp. awongkor Van Mastrigt, [1988]
ssp. hindenburgensis Lachlan, 2000

flavistriga Roepke, 1955 [endemic]
ssp. ilagaensis Van Mastrigt, [1988]
ssp. flavistriga Roepke, 1955

weiskei group (ix)

durai Van Mastrigt, 2006 [endemic]

tessei Joicey & Talbot, 1916 [endemic]
ssp. tessei Joicey & Talbot, 1916
ssp. conspectirubra Joicey & Talbot, 1922

virgo Gerrits & Van Mastrigt, [1993] [endemic]

hapalina Jordan, [1912]
ssp. adnexa Roepke, 1955
ssp. kaloni Gotts & Ginn, 2004
ssp. amoena Roepke, 1955
adjuncta Roepke, 1955
ssp. hapalina Jordan, [1912]

weiskei subgroup

leucias Jordan, [1912]
ssp. torini Gotts & Ginn, 2004
ssp. leucias Jordan, [1912]
lutescens Roepke, 1955
rhodea Roepke, 1955

rosamontana Roepke, 1955 [endemic]
ssp. tanakaorum Morita, 1996
ssp. osadai Yagishita, 1993
ssp. rosamontana Roepke, 1955
ssp. timur Van Mastrigt, 2003

nieuwenhuisi Van Mastrigt, 1990
ssp. poponga Van Mastrigt, 1990

campbelli Joicey & Talbot, 1922
ssp. campbelli Joicey & Talbot, 1922
ssp. microleuca Roepke, 1955
subflavescens Roepke, 1955
ssp. cyclops Van Mastrigt, 1996

cumanau Van Mastrigt, 2006 [endemic]

niepelti group (xvi) [species in this group need a thorough study to confirm the species status]

arfakensis Joicey & Talbot, 1922

hypoxantha Roepke, 1955
insula Roepke, 1955
peninsula Roepke, 1955

neagra Roepke, 1955
ssp. albimarginata Talbot, 1929
ssp. hypochrysis Roepke, 1955
ssp. neagra Jordan, [1912]

niepelti Ribbe, 1900
ssp. henki Yagishita, 1997
ssp. niepelti Ribbe, 1900

chrysomelaena group (iii)

ladas Smith, 1894
ssp. waigeuensis Joicey & Talbot, 1917
ssp. fakfakensis Yagishita, 2003
ssp. levis Joicey & Talbot, 1922
ssp. wamenaensis Morita, 1993
ssp. yapenensis Yagishita, 1998
ssp. ladas Smith, 1894
sulfurata Roepke, 1955

nigrana group (xi)

ornytion (Godman & Salvin, 1880)
ssp. persephone Staudinger, 1895
ssp. ornytion (Godman & Salvin, 1880)

dohertyi Oberthür, 1894
ssp. dohertyi Oberthür, 1894
ssp. knowlei Joicey & Noakes, 1915

wollastoni Rothschild, 1915 [endemic]
ssp. wollastoni Rothschild, 1915
ssp. bryophila Roepke, 1955
ssp. abmisibilensis Van Mastrigt, 1990

clathrata group (xv)

neeltje Gerrits & Van Mastrigt, [1993] [endemic]

elongatus Kenrick, 1911 [endemic]
anggi Van Mastrigt, 2000

mariae subgroup

mariae Joicey & Talbot, 1916 [endemic]

menooensis Joicey & Talbot, 1922 [endemic]
ssp. menooensis Joicey & Talbot, 1922
ssp. boschmai Roepke, 1955

bobaga Van Mastrigt, 1990 [endemic]
ssp. bobaga Van Mastrigt, 1990
ssp. homeyo Van Mastrigt, 1996

sigit Van Mastrigt, 1990 [endemic]

walshae Roepke, 1955 [endemic]
ssp. ilu Van Mastrigt, 2000
ssp. walshae Roepke, 1955
ssp. sanaeae Sakuma, 1999

hemianops Gerrits & Van Mastrigt, [1993]

fioretti Van Mastrigt, 1996 [endemic]

catocausta Jordan, [1912] [endemic]
ssp. eefi Van Mastrigt, 1990
ssp. catocausta Jordan, [1912]

mira subgroup

roepkei Sanford & Bennett, 1955
ssp. cieko Arima, 1996
ssp. reissingeri Van Mastrigt, 1998

inexpectata Rothschild, 1915 [endemic]

hiemalis Roepke, 1955 [endemic]
ssp. flabella Van Mastrigt, 1996
ssp. hiemalis Roepke, 1955
ssp. labbei Van Mastrigt, 2000
ssp. nemangkawi Van Mastrigt, 2000

autumnalis Roepke, 1955 [endemic]
ssp. hiberna Van Mastrigt, 2000
ssp. autumnalis Roepke, 1955
ssp. michiae Nakano, 1994

nakanokeikoae Yagashita, 1993 [endemic]
ssp. nakanokeikoae Yagashita, 1993
ssp. jali Van Mastrigt, 1996

albertisi group (xiv)

albertisi (Oberthür, 1880) [endemic]
ssp. albertisi (Oberthür, 1880)
ssp. albiplaga Joicey & Talbot, 1922
ssp. tamamitsui Morita, 1996
ssp. putih Van Mastrigt, 1996

discus Honrath, 1886
ssp. discus Honrath, 1886
ssp. larseni Lück& Gehlen, 1911
ssp. apodiscus Roepke, 1955
ssp. discoides Talbot, 1937

telefominensis Yagishita, 1993
ssp. telefominensis Yagishita, 1993
ssp. ayamiae Sakuma, 1996 comb. nov.

bornemanni group (vii)

caroli Kenrick, 1909 [endemic]
ssp. caroli Kenrick, 1909
flava Joicey & Noakes, 1915
ssp. wandammenensae Joicey & Talbot, 1916

thompsoni Joicey & Talbot, 1916 [endemic]

nais Jordan, [1912]
ssp. denigrata Joicey & Talbot, 1927
ssp. maruyamai Yagishita, 1993
ssp. odilae Gotts & Ginn, 2004
ssp. holophaea Roepke, 1955
ssp. nais Jordan, [1912]
zebra Yagishita, 1993
ssp. rubrina Van Eecke, 1915

zebra Roepke, 1955 [endemic]
ssp. takanamii Yagishita, 1993
ssp. zebra Roepke, 1955
reducta Roepke, 1955

kristianiae Van Mastrigt, 2006 [endemic]

klossi Rothschild, 1915 [endemic]
ssp. klossi Rothschild, 1915
ssp. gome Van Mastrigt, 2000
ssp. chrysanthemum Roepke, 1955
deluna Van Mastrigt, 2000
ssp. okse Van Mastrigt, 2000

pratti Kenrick, 1909 [endemic]

castaneus Kenrick, 1909 [endemic]
denigrata Küppers, 1978
atrata Küppers, 1978

ormoensis Van Mastrigt, 2006 [endemic]

hypomelas Rothschild & Jordan, 1907
ssp. rubrostriata JOICEY & TALBOT, 1922
ssp. fulgida ROEPKE, 1955
ssp. lieftincki ROEPKE, 1955
ssp. conversa JORDAN, [1912]

argentata Roepke, 1955 [endemic]
ssp. clutus Yagishita, 1993
ssp. argentata Roepke, 1955
sanguinea Roepke, 1955

takashii Sakuma, 1999 [endemic] (probably a form of Delias argentata or a hybrid)

destrigata Van Mastrigt, 1996 [endemic] (probably a form of argentata)

heroni Kenrick, 1909 [endemic]
albo-oculatus Joicey & Noakes, 1915

sphenodiscus Roepke, 1955 [endemic]

 

(Checklist last updated on 9th February 2009)

Literature

Davenport, Chr. & H. van Mastrigt, 2008. Revision of Delias mysis (Fabricius, 1775) and closely related species (Lepidoptera: Pieridae). Suara Serangga Papua (Sugapa) 3(2): 15-23.
Mastrigt, H. van, 2006. New (sub)species of Delias (Hübner, 1819) from the Foja Mountains, Papua, Indonesia (Lepidoptera: Pieridae). Futao 51: 11-18.
Parsons, M., 1999. The butterflies of Papua New Guinea. Their systematics and biology. 736 pp., Academic Press, London.